The Primary Couple in a Storyline of characters, there is always one couple and relationship that rises above all the others in dynamics overall.
Boomer and Lizbeth For The M.O.D.A.B Storyline as while they weren't a couple until later on the story and had another love interest. They had the most focus and change in that respect, starting from enemies, to begurdging friends, to Boomer falling for her with her seening him as a good friend, to becoming her confidant to realzing she likes him too. With Dib and Bubbles being the Gamma Couple and Bender and Starfire being the Beta Couple to the two.
Phineas and Isabella for The B Team Storyline being the only couple to be in every story together out of all the couples. Again Bender and Starfire are the Beta Couples to them. Though there at least 2 more couples in the Multiversing Adventures that give them competition.
Lincoln & Lydia for Phase 4 in general and The Team Free Will Storyline. Their relationship and it's development is a central focus of the Phase 4 storyline. Lincoln and Lydia are best friends and partners but throughout Return of The Weirdmagedon Cult, the two developed a crush on each other respectfully. Then this mutual crush grew into a realization that both of them are each other's one true love in To Live and/or Die In Loch Loud when Lincoln opens up to Lydia about his frustrations with everything and everyone as she is the only person he can confide in and Lydia for her part decides to forgo killing Morag immediately to get him back on his feet and help deal with his trauma. While Lincoln does have some attachment and feelings for Hilda, Lincoln only did so to distract them his love for Lydia as he was afraid to lose her if he confessed to her straight out or if something happened, Lydia worries on her end about ruining her friendship with Lincoln if she does. Team Free Will: Diassembled sees them both trying to get the courage to confess their love to each other while having some romantic moments with one another, though this in intruded on by others, usually Hilda who unlike the rest of Team Free Will doesn't know about the "thing" going on between them. At the end, Lydia kisses Lincoln on the lips randomly and she freaks out as she wasn't trying to do so and just wanted to try to thank him for helping her. Lincoln is gentle and states that he proably would did a similar thing in Loch Loud after Lydia had him pull himself together. They do manage to become a couple in the halfway point of Countdown and decide to keep them between them to deny the satisfcation of their friends teasing them about their relationship.
Fox and Misty can be highlighted throughout Fanfictiondreamer's creation, The Ultimate Story. What started out as close friends as Misty was practicing her princess/fairy training in Corneria for a full year, Fox always encouraged Misty to do her best, as he was by Lisa's side as she created the Academy starting out as a fairy. Their relationship grew though they didn't acknowledge it, but Peach and Slippy knew. By the time it was uncovered in US2. Fox denied it at first, but his friends explained Misty was the one who gave Fox hope after all his tragic losses like James and Pikachu. And he was the one who gave Misty encouragement thru her hard work and adventures. So, eventually they became an official wedded couple as well.
Ratchet and Kim Possible had quite a development throughout the Ratchet and Kim Possible Chronicles, even at the start, Kim Possible: The Extremely Secret Files. FFD loves them so much, she came up with a duet, "----"
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